2015 – 2016
Habilitation at the University of Bern in Biological Systematics

2005 – 2010
PhD at the Institute of Ecology and Evolution of the University of Bern: 'Taxonomy, phylogeny, and the evolution of courtship behaviour in Diplazontinae parasitoid wasps (Hymenoptera, Ichneumonidae)'

2004 – 2005
Master in Biology (Zoology) at the University of Bern: 'The fate of mutations surfing on the wave of a range expansion'


Since 2018
Curator of Entomology at the Natural History Museum Basel

2015 – 2018
Group leader under a grant of the Swiss National Science Foundation ('Ambizione') at the Natural History Museum in Bern and the University of Bern

2013 – 2015
Postdoc at the University of Adelaide in Australia

2010 – 2013
Postdoc at the Naturhistoriska Riksmuseet in Stockholm Sweden


Diversity, phylogeny and evolution of ichneumonid parasitoid wasps (Hymenoptera, Ichneumonidae). Calibrating trees using fossils. Co-evolution and its impact on diversification.


Klopfstein, S., Spasojevic, T. (preprint): Illustrating phylogenetic placement of fossils using RoguePlots: An example from ichneumonid parasitoid wasps (Hymenoptera, Ichneumonidae) and an extensive morphological matrix.

Klopfstein, S., Langille, B., Spasojevic, T., Broad, G.R., Cooper, S.J.B., Austin, A.D., Niehuis, O. (in press): Hybrid capture data unravels a rapid radiation of pimpliform parasitoid wasps (Hymenoptera: Ichneumonidae: Pimpliformes). Systematic Entomology.

Korenko, S., Spasojevic, T., Pekár, S., Walter, G.H., Korenková, V., Hamouzová, K., Kolářová, M., Kysilková, K., Klopfstein, S. (2018): One generalist or several specialist species? Wide host range and diverse manipulations of the hosts’ web building behaviour in the true spider parasitoid Zatypota kauros (Hymenoptera: Ichneumonidae). Insect Cons Div.

Kopylov, D.M., Spasojevic, T., Klopfstein, S. (2018): New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East. Zootaxa. 4442(2): 319–330.

Spasojevic, T., Wedemann, S., Klopfstein, S. (2018): Seven remarkable new fossil species of parasitoid wasps (Hymenoptera, Ichneumonidae) from the Eocene Messel Pit. PLoS One 13(6): e0197477.

Klopfstein, S., Van der Schyff, G., Tierney, S., Austin, A. (2018): Wolbachia infections in Australian ichneumonid parasitoid wasps (Hymenoptera: Ichneumonidae): evidence for adherence to the global equilibrium hypothesis. Biol J Linn Soc 123(3): 518–534.

Spasojevic, T., Broad, G., Bennett, A., Klopfstein, S. (2018): Ichneumonid parasitoid wasps from the Early Eocene Green River Formation: five new species and a revision of the known fauna (Hymenoptera, Ichneumonidae). Paläontologische Zeitschrift 92(1): 35-63.

Further publications on Google Scholar


Diversity of Ichneumonidae in Switzerland

Phylogeny and evolution of ichneumonid parasitoid wasps

Fossil ichneumonids and their placement in the tree of extant taxa

Calibrating molecular trees with fossils